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7.1.27 · Transport in Plants

Xylem and Phloem Distribution — Cambridge AS & A Level Biology 9700 Revision Notes & Topical Questions

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7.1.2 Xylem and Phloem Distribution — CAMBRIDGE AS & A LEVEL BIOLOGY 9700 — REVISION NOTES notes: Describe where xylem and phloem lie in roots, stems and leaves of herbaceous dicots and relate their positions to support, transport and resistance to stress.

  • Vascular bundles
  • Water entry
  • Woody plants
  • Annual rings
  • Orientation
  • Short paths

Part 1 of 8Two transport tissues

Xylem
  • Xylem it carries dissolved minerals and water up the plant from the roots and also provides structural support.
Phloem
  • Phloem it carries organic solutes, mainly sucrose and amino acids (assimilates), from sources to sinks.
Vascular bundles
  • Vascular bundles the two tissues run together in bundles that connect the whole plant, from root tips to young leaves.
Position
  • Position the location of bundles depends on the organ, because different organs are under different stresses.

Part 2 of 8Distribution in the root

1Layout
  • Layout the xylem forms a central star or X, with phloem in the gaps between its arms.
2Strength
  • Strength the central xylem lets the root resist the tension forces when the shoot is blown about, like a flexible, strong rope.
3Water entry
  • Water entry the xylem arms reach close to the endodermis, so water crossing the cortex enters the xylem after a short distance.
4Pericycle
  • Pericycle it surrounds the vascular tissue and gives rise to lateral roots.

Part 3 of 8Distribution in the stem

Layout
  • Layout the vascular bundles form a ring near the outside, with xylem on the inner side, phloem on the outer side and cambium between.
    • (closest to the epidermis)
Strength
  • Strength strong lignified xylem at the periphery resists bending best, as a tube is stiffer than a solid rod of the same mass.
Woody plants
  • Woody plants the cambium produces secondary xylem (wood) each year inward and secondary phloem outward, so the stem thickens.
Annual rings
  • Annual rings they reflect seasonal differences in vessel size in spring and summer,
    • which allows a tree to be dated, with the oldest wood in the centre.

Part 4 of 8Distribution in the leaf

Layout
  • Layout the midrib has a large vascular bundle, with smaller veins branching through the blade.
Orientation
  • Orientation xylem is on the upper side and phloem on the lower side of each bundle.
    • (nearest the lower epidermis)
Short paths
  • Short paths veins are close to every mesophyll cell, so water has a short distance from xylem to cells and sucrose has a short path to the phloem.
Support
  • Support the network of veins also supports the thin blade.
OrganXylemPhloem
RootCentral, X-shapedBetween xylem arms
StemInner side of each bundleOuter side of each bundle
LeafUpper side of bundleLower side of bundle

Part 5 of 8Identifying the tissues on a slide

Xylem
  • Xylem large, empty, thick-walled cells with no contents, whose lignin is easy to see when stained.
Phloem
  • Phloem smaller cells, with thin-walled sieve tube elements and companion cells with dense cytoplasm and a nucleus placed next to them.
Sieve plates
  • Sieve plates they are sometimes visible in a transverse section.

Part 6 of 8Evidence for the direction of transport

Ringing
  • Ringing removing a ring of bark including phloem makes sugars accumulate above the ring, which shows that phloem carries organic solutes downward.
  • The xylem inside is unaffected, so water still moves upward.
Radioactive tracers
  • 14C in carbon dioxide can be used to follow assimilates in the phloem.
Aphid stylets
  • Aphid stylets they can be used to sample phloem sap.

Part 7 of 8Support, lignin and turgor

Support, lignin and turgor
Turgor
  • Turgor water-filled cells press outward against their cellulose walls and keep herbaceous stems rigid,
    • and loss of water lowers turgor so the plant wilts.
Lignified tissues
  • Lignified tissues lignified xylem and thickened collenchyma and sclerenchyma add strength.
Position and function
  • Position and function xylem near the centre of a root resists pulling, peripheral stem bundles resist bending,
    • and leaf veins sit close to the mesophyll for short diffusion paths.

Part 8 of 8More from the specification

More from the specification
Plant Vascular Bundles and Tissue Distribution
  • In dicotyledonous roots, xylem forms a central X-shaped core surrounded by phloem;
    • in stems, vascular bundles are arranged in a peripheral ring with xylem internal to phloem.
Structure and Function of Mature Xylem Vessels
  • Mature xylem vessel elements are dead, hollow, elongated cells with lignified secondary walls that lack end walls,
  • forming a continuous capillary tube for water and mineral transport.
Structure of Phloem Sieve Tube Elements
  • Phloem tissue consists of living sieve tube elements with perforated end sieve plates, closely associated with metabolically active companion cells that regulate organic nutrient transport.
Micrograph Drawings and Botanical Conventions
  • Biological plan drawings made from transverse plant stem or root micrographs must represent tissue layer boundaries with clear continuous lines without drawing individual cellular details.
Micrograph
  • A photograph taken through a microscope.
Compound
  • A substance of two or more kinds of atoms chemically combined
Glucose
  • A simple sugar; the unit that starch, glycogen and cellulose are built from.
Organic compound
  • Compound containing carbon and hydrogen.
Transport system
  • A system, such as the circulatory system, that moves substances around the body.

Key terms

TermMeaning
XylemTissue that transports water and mineral ions upward and provides support.
PhloemTissue that transports assimilates such as sucrose throughout the plant.
AssimilatesOrganic substances such as sucrose and amino acids transported in the phloem.
SourceA region where assimilates are loaded into the phloem, such as a photosynthesising leaf.
SinkA region where assimilates are removed from the phloem and used or stored.
CambiumA layer of dividing cells between xylem and phloem that produces new vascular tissue.
PericycleA layer of cells inside the endodermis of a root that can produce lateral roots.

Avoid these mistakes

1
Reversing the positions of xylem and phloem in the stem bundle.
2
Saying phloem carries water upward — xylem carries water up and phloem carries assimilates in either direction.
3
Saying only xylem is dead — phloem sieve tubes are living cells.
4
Forgetting the cambium between xylem and phloem in a stem bundle.
5
Stating that xylem carries only water — it carries mineral ions as well.

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