7.1.27 · Transport in Plants
Xylem and Phloem Distribution — Cambridge AS & A Level Biology 9700 Revision Notes & Topical Questions
5 min read 10 parts Lesson 4 of 15 1 quiz
Lesson summary
7.1.2 Xylem and Phloem Distribution — CAMBRIDGE AS & A LEVEL BIOLOGY 9700 — REVISION NOTES notes: Describe where xylem and phloem lie in roots, stems and leaves of herbaceous dicots and relate their positions to support, transport and resistance to stress.
- Vascular bundles
- Water entry
- Woody plants
- Annual rings
- Orientation
- Short paths
Part 1 of 8Two transport tissues
Xylem
- Xylem it carries dissolved minerals and water up the plant from the roots and also provides structural support.
Phloem
- Phloem it carries organic solutes, mainly sucrose and amino acids (assimilates), from sources to sinks.
Vascular bundles
- Vascular bundles the two tissues run together in bundles that connect the whole plant, from root tips to young leaves.
Position
- Position the location of bundles depends on the organ, because different organs are under different stresses.
Part 2 of 8Distribution in the root
1Layout
- Layout the xylem forms a central star or X, with phloem in the gaps between its arms.
2Strength
- Strength the central xylem lets the root resist the tension forces when the shoot is blown about, like a flexible, strong rope.
3Water entry
- Water entry the xylem arms reach close to the endodermis, so water crossing the cortex enters the xylem after a short distance.
4Pericycle
- Pericycle it surrounds the vascular tissue and gives rise to lateral roots.
Part 3 of 8Distribution in the stem
Layout
- Layout the vascular bundles form a ring near the outside, with xylem on the inner side, phloem on the outer side and cambium between.
- (closest to the epidermis)
Strength
- Strength strong lignified xylem at the periphery resists bending best, as a tube is stiffer than a solid rod of the same mass.
Woody plants
- Woody plants the cambium produces secondary xylem (wood) each year inward and secondary phloem outward, so the stem thickens.
Annual rings
- Annual rings they reflect seasonal differences in vessel size in spring and summer,
- which allows a tree to be dated, with the oldest wood in the centre.
Part 4 of 8Distribution in the leaf
Layout
- Layout the midrib has a large vascular bundle, with smaller veins branching through the blade.
Orientation
- Orientation xylem is on the upper side and phloem on the lower side of each bundle.
- (nearest the lower epidermis)
Short paths
- Short paths veins are close to every mesophyll cell, so water has a short distance from xylem to cells and sucrose has a short path to the phloem.
Support
- Support the network of veins also supports the thin blade.
| Organ | Xylem | Phloem |
|---|---|---|
| Root | Central, X-shaped | Between xylem arms |
| Stem | Inner side of each bundle | Outer side of each bundle |
| Leaf | Upper side of bundle | Lower side of bundle |
Part 5 of 8Identifying the tissues on a slide
Xylem
- Xylem large, empty, thick-walled cells with no contents, whose lignin is easy to see when stained.
Phloem
- Phloem smaller cells, with thin-walled sieve tube elements and companion cells with dense cytoplasm and a nucleus placed next to them.
Sieve plates
- Sieve plates they are sometimes visible in a transverse section.
Part 6 of 8Evidence for the direction of transport
Ringing
- Ringing removing a ring of bark including phloem makes sugars accumulate above the ring, which shows that phloem carries organic solutes downward.
- The xylem inside is unaffected, so water still moves upward.
Radioactive tracers
- 14C in carbon dioxide can be used to follow assimilates in the phloem.
Aphid stylets
- Aphid stylets they can be used to sample phloem sap.
Part 7 of 8Support, lignin and turgor
Support, lignin and turgor
Turgor
- Turgor water-filled cells press outward against their cellulose walls and keep herbaceous stems rigid,
- and loss of water lowers turgor so the plant wilts.
Lignified tissues
- Lignified tissues lignified xylem and thickened collenchyma and sclerenchyma add strength.
Position and function
- Position and function xylem near the centre of a root resists pulling, peripheral stem bundles resist bending,
- and leaf veins sit close to the mesophyll for short diffusion paths.
Part 8 of 8More from the specification
More from the specification
Plant Vascular Bundles and Tissue Distribution
- In dicotyledonous roots, xylem forms a central X-shaped core surrounded by phloem;
- in stems, vascular bundles are arranged in a peripheral ring with xylem internal to phloem.
Structure and Function of Mature Xylem Vessels
- Mature xylem vessel elements are dead, hollow, elongated cells with lignified secondary walls that lack end walls,
- forming a continuous capillary tube for water and mineral transport.
Structure of Phloem Sieve Tube Elements
- Phloem tissue consists of living sieve tube elements with perforated end sieve plates, closely associated with metabolically active companion cells that regulate organic nutrient transport.
Micrograph Drawings and Botanical Conventions
- Biological plan drawings made from transverse plant stem or root micrographs must represent tissue layer boundaries with clear continuous lines without drawing individual cellular details.
Micrograph
- A photograph taken through a microscope.
Compound
- A substance of two or more kinds of atoms chemically combined
Glucose
- A simple sugar; the unit that starch, glycogen and cellulose are built from.
Organic compound
- Compound containing carbon and hydrogen.
Transport system
- A system, such as the circulatory system, that moves substances around the body.
Key terms
| Term | Meaning |
|---|---|
| Xylem | Tissue that transports water and mineral ions upward and provides support. |
| Phloem | Tissue that transports assimilates such as sucrose throughout the plant. |
| Assimilates | Organic substances such as sucrose and amino acids transported in the phloem. |
| Source | A region where assimilates are loaded into the phloem, such as a photosynthesising leaf. |
| Sink | A region where assimilates are removed from the phloem and used or stored. |
| Cambium | A layer of dividing cells between xylem and phloem that produces new vascular tissue. |
| Pericycle | A layer of cells inside the endodermis of a root that can produce lateral roots. |
Avoid these mistakes
1
Reversing the positions of xylem and phloem in the stem bundle.
2
Saying phloem carries water upward — xylem carries water up and phloem carries assimilates in either direction.
3
Saying only xylem is dead — phloem sieve tubes are living cells.
4
Forgetting the cambium between xylem and phloem in a stem bundle.
5
Stating that xylem carries only water — it carries mineral ions as well.
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