4.4.5Unit 4: Energy, Environment, Microbiology and Immunity
Index of Biodiversity — IAL Edexcel Linear Revision Notes & Topical Questions
5 min read 5 parts Lesson 25 of 28 Unit quiz
Part 1 of 5The Index

An index of diversity links the number of species to the number of individuals in each, so it uses richness and abundance together.
Formula
- D = N(N − 1) divided by the sum of n(n − 1) for every species in the community.
N
- The total number of organisms of all species in the community.
n
- The number of individuals of a single species.
Interpretation
- A larger value of D means a higher level of diversity.
Use
- Calculating D for different habitats lets their diversity be compared, or one habitat compared over time.
Part 2 of 5Calculation Steps
Work in the same order each time and show each value.
1
Count all the organisms of all species to give N, then work out N(N − 1) as value A.
2
For each species calculate n(n − 1), then add the results together to give value B.
3
Divide A by B to obtain the index of diversity D and give it to the required decimal places.
4
Compare D for different habitats: the higher value shows the more diverse habitat.
Part 3 of 5Why Use an Index
Why Use an Index
Evenness
- If one species dominates, the denominator becomes large and D falls, correctly showing lower diversity.
Rare species
- Rare species contribute little to the denominator, so a community with many evenly spread species gets a high D.
Minimum value
- If all individuals are of one species, A and B are equal and D is 1, the lowest possible value.
Sampling
- Results depend on sampling method and effort, so the same method should be used for every habitat compared.
Part 4 of 5Worked Example
Question
- A sample of insects from a hedge contains 12, 8, 5, 3 and 2 individuals of five species.
- Calculate the index of diversity.
- N = 12 + 8 + 5 + 3 + 2 = 30, so A = N(N − 1) = 30 × 29 = 870.
- n(n − 1): 12×11 = 132; 8×7 = 56; 5×4 = 20; 3×2 = 6; 2×1 = 2. Sum B = 216.
- D = 870 ÷ 216 = 4.03 (to 2 decimal places).
Answer D = 4.03
Part 5 of 5Exam Checklist & Tips
4.4.5 – Key Points for the Exam
- Index of diversity A value that combines the number of species and their relative abundance.
- Abundance The number of individuals of a species in an area.
- Evenness How equally individuals are spread among the species present.
- Sigma (Σ) The symbol meaning the sum of all the values that follow.
Common Mistakes
- Using the number of species in place of the total number of organisms as N.
- Forgetting to calculate n(n − 1) for every species before adding them together.
- Dividing in the wrong order, giving Σn(n − 1) ÷ N(N − 1) instead of the other way round.
- Concluding a habitat is diverse from a single high D value without comparing to another habitat.
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