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5.1Topic 5 · On the Wild Side

Ecosystems and Energy Transfer — Edexcel A Level Biology A (SNAB) 9BI0 Revision Notes & Topical Questions

15 min read 25 parts Lesson 3 of 34 Unit quiz

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Ecosystems, Habitats and Niches — EDEXCEL A LEVEL BIOLOGY A (SNAB) 9BI0 — REVISION NOTES notes: Define ecosystem terms and explain how biotic and abiotic factors and niches determine abundance and distribution.Species are adapted to life in a particular habitat.

  • population
  • Worked example 1
  • Worked example 2
  • Worked example 3
  • Saying a habitat and an ecosystem are the same
  • Listing predation as abiotic

Section 1 of 25Habitat, population, community and ecosystem

Species are adapted to life in a particular habitat.

Habitat, population, community and ecosystem
Habitat
A habitat is the place where an organism lives; it can be large (a desert) or small (an individual tree).
Population
Members of a species in a habitat form a population; populations interact in a community;
  • and communities interact with each other and with the non-living parts of their habitat to form an ecosystem.
An ecosystem is a community and its interactions with the non-living parts of its habitat.
Energy flows through an ecosystem and nutrients are recycled within it.
Ecosystems vary greatly in size and scale: a small pond in a garden and the open ocean can both be called ecosystems.

Section 2 of 25Biotic and abiotic factors

The abundance and distribution of a species in a habitat are determined by a combination of biotic and abiotic factors. Biotic factors are living factors that result from the activity of other organisms: intraspecific competition (individuals of the same species competing for resources), interspecific competition (between different species), predation, disease, parasitism and food availability. Abiotic factors are non-living factors, such as temperature, light intensity, pH, soil moisture, salinity, oxygen or carbon dioxide concentration and wind. For example, low light reduces plant growth in a woodland floor, and a low oxygen concentration in a polluted river favours organisms that tolerate it.

Abiotic factorImpact on a community
TemperatureAffects enzyme activity, rates of reactions, growth and breeding
Light intensityLimits photosynthesis and so producer growth
pHAffects enzymes and nutrient availability in soil
Oxygen concentrationLimits aerobic organisms in water

Section 3 of 25Niches

Species occupy a specific niche in a habitat.

Niches
A niche is the role of a species within its habitat,
  • including what it eats,
  • where it feeds,
  • when it is active and its interactions with other species and with abiotic factors.
Features such as feeding location may differ between niches, so closely related species can coexist.
Two species cannot occupy exactly the same niche for long: competition would cause one to be excluded (competitive exclusion).

Section 4 of 25Measuring distribution and abundance

The abundance and distribution of species can be found by counting all the organisms in a habitat if the area is small, but usually sampling is used (see the next lesson).

Section 5 of 25Interactions within communities

Intraspecific competition occurs when members of one species compete for the same limited resources such as food, mates and space, and it tends to control population size.

Interactions within communities
Interspecific competition occurs between species with overlapping niches; the better competitor may eliminate the other from that habitat (competitive exclusion).
Predator-prey populations often fluctuate in cycles: as prey increase, predators increase, which then reduces prey, and then predator numbers fall.
These interactions help determine where and how many individuals of a species are found.

Section 6 of 25Worked examples

  1. Distinguish between a habitat and a niche. [2]2 marks

  2. Give two examples of biotic factors affecting a population. [2]2 marks

  3. Define ecosystem. [2]2 marks

Section 7 of 25Random and systematic sampling

Random sampling selects sample sites at random, for example by using a random number table to generate co-ordinates on a grid laid over the habitat and placing quadrats at those points. This avoids bias and is suitable for areas that are uniform. Systematic sampling places sample sites at regular intervals, and is used to investigate a change along a gradient.

Section 8 of 25Quadrats

A frame quadrat is a square frame (commonly 0.25 m2 or 1 m2).

Species frequency
Depending on the aim, record species frequency (how many quadrats contain the species or how many individuals are in the quadrat) or abundance estimated by percentage cover.
Point quadrat
A point quadrat is a vertical frame with holes across the top through which pins are lowered; each species touched by a pin is recorded.
It is useful in dense plant cover where the ground is hard to study with a frame quadrat and can measure the number of individuals of a species or percentage cover.

Section 9 of 25Transects

Transects are lines laid across a site to measure abundance and distribution across a habitat, useful for determining how species abundance and distribution change along a gradient, for example at increasing distance from a field margin or perpendicular to the water's edge on a rocky shore.

Line transect
A line transect records species touching the line; a belt transect uses a strip of a set width; an interrupted transect samples at intervals along the line, often with quadrats.
The distribution and abundance of species are affected by abiotic factors,
  • so when investigating the impact of one, measure the relevant abiotic factors at each sample site.
    • (light, pH, soil moisture, temperature)

Section 10 of 25Kite diagrams

Results can be shown on a kite diagram.

Kite diagrams
The distribution of a species along a transect is shown by its position along a central horizontal line,
  • and its abundance by the width of the kite at each point.
Additional sections can show abiotic changes such as height above sea level or soil pH.
Kite diagrams therefore show both abundance and distribution of species and changes in abiotic factors.

Planning tipwhen asked to describe or design an investigation, include what you will vary, how you will measure the dependent variable (e.g. percentage cover), how you will ensure validity (control other variables, random placement, enough repeats) and the choice of technique and why.

Section 11 of 25Choosing the right method

Use a frame quadrat for plants or slow-moving animals in a small area, and random sampling in a uniform habitat.

1
Use a transect with systematic sampling where there is a gradient such as a rocky shore, a sand dune or distance from a path.
2
Use a point quadrat for tall dense grassland.
3
For animals that move, use pitfall traps, sweep nets or mark-release-recapture.
4
Reliability improves with more samples;
  • and the validity of conclusions about the cause requires abiotic factors to be measured and compared, for example using a statistical test such as Spearman's rank correlation.

Section 12 of 25Worked examples

  1. Explain why random sampling is used. [2]2 marks

  2. Which technique would you use to investigate how species change from the edge of a pond to dry land? [2]2 marks

  3. What does the width of a kite diagram show? [1]1 mark

Section 13 of 25Succession

Ecosystems are dynamic

  • they change constantly.
  • The process of ecosystem change over time is succession, during which biotic and abiotic conditions change. Primary succession starts on bare ground such as rock or sand dunes, with no soil;
    • secondary succession occurs where soil already exists, for example after a fire or abandoned farmland.

Section 14 of 25Stages of primary succession

1Pioneer species
Pioneer species such as lichens and algae colonise bare rock.
  • They break apart the top surface;
    • this fragmented rock, together with dead organic matter from dead lichens, forms a basic soil.
2
Species such as mosses and grasses grow, and their roots stabilise the soil, enabling it to hold more moisture and nutrients.
  • As they die, humus builds up.
3
Larger plants and shrubs, as well as small trees, can now grow. These need more water, which can be stored in deeper soils.
4Dominant species
Over time the soil becomes deep enough, with enough nutrients and water, to support large trees.
  • The final species to colonise become the dominant species of the now complex climax community, which is relatively stable.

Often the new colonising species change the environment so that it becomes less suitable for the previous species.

1
For example, lichens cannot grow on soil, so they disappear once soil forms.
2
As succession proceeds, species diversity and biomass increase, food webs become more complex, and abiotic conditions become less extreme.
3
Be able to apply your knowledge to any example, such as sand dunes or a newly formed volcanic island.

Section 15 of 25Net primary productivity

In photosynthesis, producers convert light energy into chemical energy stored in biological molecules.

1Gross primary productivity (GPP)
The rate at which producers do this is gross primary productivity (GPP).
2Net primary productivity (NPP)
Some of this energy is used in respiration (R) and lost as heat. Net primary productivity (NPP) is the rate at which energy is stored in plant biomass and made available to primary consumers: NPP = GPP - R.
3
Rearranged: GPP = NPP + R, and R = GPP - NPP.
4
Units are usually kJ m-2 yr-1 (energy per area per time).

Worked example

  • grass in a meadow converts light energy to carbohydrate at a rate of 17 500 kJ m-2 yr-1 (GPP) and releases 14 000 kJ m-2 yr-1 by respiration (R).
  • NPP = 17 500 - 14 000 = 3 500 kJ m-2 yr-1.

Section 16 of 25Succession in sand dunes

On a sand dune system, the youngest dunes near the sea are colonised by pioneer species such as marram grass, which tolerates salt, wind and low water, and whose roots bind the sand.

Succession in sand dunes
As humus accumulates, soil nutrients and water retention increase,
  • so that fescue grasses,
  • then shrubs, and finally woodland develop.
    • (in older dunes further inland)
Abiotic factors such as pH, humus content and soil moisture can be measured along a transect inland to show the gradient of succession.

Section 17 of 25Worked examples

  1. Explain why lichens disappear during succession. [3]3 marks

  2. GPP is 12 000 and NPP is 4 500 kJ m−2 yr-1. Calculate R. [2]2 marks

  3. Describe how soil forms during primary succession. [3]3 marks

Section 18 of 25Energy flow and losses

In photosynthesis, plants convert light energy into chemical energy in biological molecules.

Energy flow and losses
Energy is then passed along food chains, but energy transfer is not 100% efficient: energy is lost to the environment at every trophic level.
A large proportion of the sun's energy is not available to producers for building biomass: some light passes through leaves or is reflected, some is the wrong wavelength,
  • and some falls on non-photosynthetic parts.

Losses between trophic levels include

  • not all of an organism is eaten (bones, roots, shells);
    • not all that is eaten is digested and absorbed (lost in faeces);
    • energy is lost as heat through respiration;
    • energy is lost in excretion (urine) and movement.

Energy flow and losses
Only the remaining energy is stored in new biomass available to the next level.
This explains why food chains are short, typically with four or five trophic levels.

Section 19 of 25Calculating efficiency of energy transfer

Energy efficiency = (net productivity of the trophic level / energy received from the previous level) x 100.

Worked examplea toad receives 10 000 kJ m-2 yr-1 and has a net productivity of 1 000 kJ m-2 yr-1.

Calculating efficiency of energy transfer
Efficiency = (1 000 / 10 000) x 100 = 10%.
Typical efficiencies are about 10% from one trophic level to the next, though they are higher for herbivores to carnivores than plants to herbivores in some cases.

Section 20 of 25Calculating efficiency of biomass transfer

Efficiency of biomass transfer = (biomass transferred / biomass intake) x 100. Check that both quantities are in the same units; if both are in kg no conversion is needed.

Worked example

  • a consumer ingests 35 kg of biomass and 4.1 kg is incorporated into its body.
  • Efficiency = (4.1 / 35) x 100 = 11.7%, which is 12% to two significant figures.

Section 21 of 25Farming implications

Farmers increase the efficiency of food production by reducing energy losses: keeping animals indoors (less movement and heat loss), controlling parasites and pests, and feeding crops directly to humans rather than to livestock, since eating at a lower trophic level wastes less energy.

Section 22 of 25Why this matters for food production

Because only about 10% of energy is transferred at each step, a given area of land can support far more people if they eat plants directly than if the plants are fed to animals first.

Why this matters for food production
Farmers raise the efficiency of livestock production by restricting movement, keeping animals warm and preventing disease,
  • which cuts energy lost in respiration and parasitism.
Efficiency can also be improved with fast-growing breeds.
These measures raise ethical questions about animal welfare which must be weighed against efficiency.

Section 23 of 25Worked examples

  1. A herbivore receives 25 000 kJ and stores 2 500 kJ in new biomass. Calculate efficiency. [2]2 marks

  2. Give three reasons for energy loss between trophic levels. [3]3 marks

  3. Explain why food chains rarely have more than five trophic levels. [2]2 marks

Section 24 of 25Key terms

TermMeaning
HabitatPlace where an organism lives.
PopulationMembers of one species in a habitat.
CommunityAll populations in a habitat.
NicheRole of a species in its habitat.
Abiotic factorNon-living factor.
Biotic factorLiving factor.
TermMeaning
Frame quadratSquare frame used to sample plants.
Point quadratFrame with pins lowered to record species touched.
TransectLine across a habitat used for sampling.
Kite diagramGraph showing abundance and distribution along a transect.
Percentage coverProportion of the quadrat covered by a species.
TermMeaning
SuccessionChange in an ecosystem over time.
Pioneer speciesFirst species to colonise bare ground.
Climax communityStable community at the end of succession.
Gross primary productivityRate of energy capture by producers.
Net primary productivityGPP minus energy used in respiration.
TermMeaning
Trophic levelPosition of an organism in a food chain.
BiomassMass of living material.
Efficiency of transferPercentage of energy passed to the next level.
Net productivityEnergy stored in new biomass.

Section 25 of 25Exam technique

1
Saying a habitat and an ecosystem are the same.
2
Listing predation as abiotic.
3
Saying that intraspecific competition is between different species.
4
Thinking that a niche is a place.
1
Using random sampling along a gradient.
2
Failing to measure abiotic factors at each site.
3
Placing quadrats at the sites that look most interesting.
4
Reading a kite diagram width as the length of the transect.
1
Saying NPP = GPP + R.
2
Saying succession ends with no further change at all.
3
Saying that primary succession starts with soil.
4
Writing that NPP is the energy captured by the producers in photosynthesis, which is GPP.
1
Saying energy is destroyed — it is transferred to the environment as heat.
2
Using the wrong denominator in the efficiency formula.
3
Saying energy is recycled in ecosystems — it flows through, while nutrients are recycled.
4
Quoting the efficiency as a number without a percentage sign or units.

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