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4.11.3Additional lessons

Keystone Species — IB Biology HL Revision Notes & Topical Questions

5 min read 5 parts Lesson 94 of 101

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Part 1 of 5What Makes a Species Keystone

4.11.3 Keystone Species

Keystone species hold a community together in the way a keystone holds an arch.

Definition
  • A keystone species has an effect on its community far larger than would be expected from its biomass or abundance.
Mechanism
  • Effects usually operate through feeding links, where the keystone controls the numbers of a dominant competitor.
Removal
  • Loss of the species can cause large changes in the community, including loss of biodiversity.
Habitat role
  • Some keystone species, such as beavers, change the physical habitat as well as the food web.
SpeciesRoleEffect of removal
Sea otterEats sea urchinsUrchins overgraze kelp forests
Pisaster ochraceus (starfish)Eats musselsMussels crowd out other species
BeaverBuilds damsWetland habitat disappears

Part 2 of 5Sea Otter Cascade

1
Sea otters feed on sea urchins, which graze on the large brown algae that form kelp forests.
2
When otters were hunted heavily, urchin numbers rose because their predator was removed.
3
Urchins overgrazed the kelp, leaving bare rocky areas called urchin barrens with few fish.
4
When otters were protected and returned, urchin numbers fell and kelp forests recovered, restoring habitat for many other species.

Part 3 of 5Identifying and Protecting Them

Identifying and Protecting Them
Removal experiments
  • Comparing plots with and without the species shows its effect on the number of other species present.
Conservation priority
  • Protecting a keystone species can protect many others at the same time, so it is cost-effective.
Trophic cascade
  • An effect that passes down several feeding levels, as with otters, urchins and kelp.
Caution
  • A species may be keystone in one ecosystem but not in another, so each case must be tested.
Examples in practice
  • In the Pacific coast intertidal zone, removing the starfish Pisaster lets mussels spread,
    • and the number of other species on the rocks falls.

Part 4 of 5Worked Example

Question

  • In a removal experiment a rocky shore plot with a starfish has 15 species and a plot without it has 6.
  • Calculate the percentage loss of species richness when the starfish is removed.

  1. Loss = 15 − 6 = 9 species.
  2. Percentage loss = 9 ÷ 15 × 100 = 60%.
  3. The large loss shows the starfish is acting as a keystone predator (values are illustrative).

Answer 60% loss of species richness

Part 5 of 5Exam Checklist & Tips

4.11.3 – Key Points for the Exam

  • Keystone species A species with a disproportionately large effect on community structure.
  • Trophic cascade A chain of effects passing through several trophic levels.
  • Overgrazing Excess feeding on plants that damages the plant community.
  • Removal experiment A field test in which a species is taken out to observe the effects.
  • Predation Feeding by one organism on another.

Common Mistakes

  • Defining keystone species as the most abundant species; abundance is not the key.
  • Forgetting to describe the whole chain of effects in a cascade.
  • Saying that all top predators are keystone species.
  • Giving the effect without linking it to a change in biodiversity.

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